Contrary to prior focus in the literature on the role of migrating infectious individuals \cite{Jesse_2008,North_2017,Grenfell_1997}, we found that the migration of immune individuals can be equally (or even more) important. This is noteworthy, as the few previous studies relating multi-strain diseases and metapopulation structure only allow pathogen transmission between populations, not the movement of individuals explicitly \cite{Lourenço2013,Wikramaratna2014}—an approach that is more mathematically tractable, but omits the potentially influential transmission of immune individuals.